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The science of ecology attempts to explain why plants and animals live where they do and why their popu- lations are the sizes they are. Understanding the distribution and population size of organisms helps scientists evaluate the health of the environment. In 1840 German chemist, Justus von Liebig first proposed that populations could not grow indefinitely, a basic principle now known as the Law of the Minimum. Biotic and abiotic factors, singly or in combination, ultimately limit the size that any population may attain. This size limit, known as a population’s carrying capac- ity, occurs when needed resources, such as food, breeding sites, and water, are in short supply. For example, the amount of nutrients in soil influences the amount of wheat that grows on a farm. If just one soil nutrient, such as nitrogen, is missing or below optimal levels, fewer healthy wheat plants will grow. Either population size or distribution may also be affected, directly or indirectly, by the way species in an ecosystem interact with one another. In an experiment performed in the late 1960s in the rocky tidal zone along the Pacific Coast of the United States, American ecologist Robert Paine studied an area that contained 15 spe- cies of invertebrates, including starfish, mussels, limpets, barnacles, and chitons. Paine found that in this eco- system one species of starfish preyed heavily on a species of mussel, preventing that mussel population from multiplying and monopolizing space in the tidal zone. When Paine removed the starfish from the area, he found that the mussel population quickly increased in size, crowding out most other organisms from rock surfaces. The number of invertebrate species in the ecosystem soon dropped to eight species. Paine concluded that the loss of just one species, the starfish, indirectly led to the loss of an additional six species and a transformation of the ecosystem. Typically, the species that coexist in ecosystems have evolved together for many generations. These populations have established balanced interactions with each other that enable all populations in the area to remain relatively stable.